Papers available to read at: 

See Research for short abstracts



  1. Inkpen SA, Doolittle WF. 2022. Do Microbial Communities Regenerate? University of Chicago Press.



  1. Gillette K, Inkpen SA, DesRoches CT. Forthcoming. "Does Environmental Science Crowd Out Non-Epistemic Values?" Studies in History and Philosophy of Science, TBA. (For a special issue on the environmental sciences)

  2. Inkpen SA, Doolittle WF. 2021. "Adaptive Regeneration Across Scales: Replicators and Interactors from Limbs to Forests." Philosophy, Theory, and Practice in Biology, 13:1.


  1. Inkpen SA, DesRoches CT. 2020. “When Ecology Needs Economics and Economics Needs Ecology: Interdisciplinary Exchange during the Anthropocene.” Ethics, Policy & Environment, 23: 203-221.


  1. Inkpen SA. 2019. “Health, Ecology, and the Microbiome.” eLife, 8: e47626.

  2. Wideman JG, Inkpen SA, Doolittle WF, Redfield RJ. 2019. "Mutationism, not Lamarckism, captures the novelty of CRISPR-Cas" Biology and Philosophy34: 12.

  3. Inkpen SADesRoches CT. 2019. "Revamping the Image of Science for the Anthropocene." Philosophy, Theory, and Practice in Biology, 11: 1-7.

  4. DesRoches CT, Inkpen SA, Green TL. 2019. “The Eroding Artificial/Natural Distinction? Some Consequences for Ecology and Economics.” In M Nagatsu and A Ruzzene (eds.), Contemporary Philosophy and Social Science: An Interdisciplinary Dialogue. Bloomsbury Publishing, London: UK.


  1. Doolittle WF, Inkpen SA. 2018 “Processes and Patterns of Interaction as Units of Selection: An Introduction to ITSNTS Thinking” Proceedings of the National Academy of Sciences (USA), 115: 4006-4014.


  1. Inkpen SA, Douglas GM, Brunet TDP, Leuschen K, Doolittle WF, Langille MGI. 2017. “The Coupling of Taxonomy and Function in Microbiomes.” Biology and Philosophy, 32: 1225-1243.

  2. Inkpen SA. 2017. “Demarcating Nature, Defining Ecology: Creating a Rationale for the Study of Nature’s ‘Primitive Conditions’” Perspectives on Science, 25: 355-92.

  3. Inkpen SA. 2017. “Are Humans Disturbing Conditions in Ecology?” Biology and Philosophy, 32: 51-71.

  4. Stuart YE, Inkpen SA, Hopkins R, Bolnick DI. 2017. “Character displacement is an evolutionary pattern. So what causes it?” Biological Journal of the Linnean Society, 121: 711-715.


  1. Inkpen SA. 2016. “Like Hercules and the Hydra: Trade-offs and Strategies in Ecological Model-Building and Experimental Design” Studies in History and Philosophy of Biological and Biomedical Sciences, 57: 34-43.

  2. Inkpen SA, Doolittle WF. 2016. “Molecular Phylogenetics and the Perennial Problem of Homology” Journal of Molecular Evolution, 83: 184-192.


  1. Inkpen SA. 2014. “‘The Art Itself is Nature’: Darwin, Domestic Varieties, and the Scientific Revolution.” Endeavour, 38: 246-56.



  1. 2015. “Putting Nature in its Place.” RemediaNetwork, May 2015. (Online at:

  2. 2014. Denaturing Nature: Philosophical and Historical Reflections on the Artificial-Natural Distinction in the Life Sciences. Dissertation: University of British Columbia, September 2014. (Online at: