Papers available to read at: 

See Research for short abstracts



  1. Inkpen SA, Doolittle WF. In preparation (due March 2021). Do Microbial Communities Regenerate? A Philosophical Inquiry. (tentative title). University of Chicago Press.



  1. Gillette K, Inkpen SA, DesRoches CT. Forthcoming. "Does Environmental Science Crowd Out Non-Epistemic Values?" Studies in History and Philosophy of Science, TBA. (For a special issue on the environmental sciences)

  2. Inkpen SA, Doolittle WF. 2021. "Adaptive Regeneration Across Scales: Replicators and Interactors from Limbs to Forests." Philosophy, Theory, and Practice in Biology, 13:1.


  1. Inkpen SA, DesRoches CT. 2020. “When Ecology Needs Economics and Economics Needs Ecology: Interdisciplinary Exchange during the Anthropocene.” Ethics, Policy & Environment, 23: 203-221.


  1. Inkpen SA. 2019. “Health, Ecology, and the Microbiome.” eLife, 8: e47626.

  2. Wideman JG, Inkpen SA, Doolittle WF, Redfield RJ. 2019. "Mutationism, not Lamarckism, captures the novelty of CRISPR-Cas" Biology and Philosophy34: 12.

  3. Inkpen SADesRoches CT. 2019. "Revamping the Image of Science for the Anthropocene." Philosophy, Theory, and Practice in Biology, 11: 1-7.

  4. DesRoches CT, Inkpen SA, Green TL. 2019. “The Eroding Artificial/Natural Distinction? Some Consequences for Ecology and Economics.” In M Nagatsu and A Ruzzene (eds.), Contemporary Philosophy and Social Science: An Interdisciplinary Dialogue. Bloomsbury Publishing, London: UK.


  1. Doolittle WF, Inkpen SA. 2018 “Processes and Patterns of Interaction as Units of Selection: An Introduction to ITSNTS Thinking” Proceedings of the National Academy of Sciences (USA), 115: 4006-4014.


  1. Inkpen SA, Douglas GM, Brunet TDP, Leuschen K, Doolittle WF, Langille MGI. 2017. “The Coupling of Taxonomy and Function in Microbiomes.” Biology and Philosophy, 32: 1225-1243.

  2. Inkpen SA. 2017. “Demarcating Nature, Defining Ecology: Creating a Rationale for the Study of Nature’s ‘Primitive Conditions’” Perspectives on Science, 25: 355-92.

  3. Inkpen SA. 2017. “Are Humans Disturbing Conditions in Ecology?” Biology and Philosophy, 32: 51-71.

  4. Stuart YE, Inkpen SA, Hopkins R, Bolnick DI. 2017. “Character displacement is an evolutionary pattern. So what causes it?” Biological Journal of the Linnean Society, 121: 711-715.


  1. Inkpen SA. 2016. “Like Hercules and the Hydra: Trade-offs and Strategies in Ecological Model-Building and Experimental Design” Studies in History and Philosophy of Biological and Biomedical Sciences, 57: 34-43.

  2. Inkpen SA, Doolittle WF. 2016. “Molecular Phylogenetics and the Perennial Problem of Homology” Journal of Molecular Evolution, 83: 184-192.


  1. Inkpen SA. 2014. “‘The Art Itself is Nature’: Darwin, Domestic Varieties, and the Scientific Revolution.” Endeavour, 38: 246-56.



  1. 2015. “Putting Nature in its Place.” RemediaNetwork, May 2015. (Online at:

  2. 2014. Denaturing Nature: Philosophical and Historical Reflections on the Artificial-Natural Distinction in the Life Sciences. Dissertation: University of British Columbia, September 2014. (Online at: